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Alan Knapp

Alan Knapp

Professor, Department of Biology, Colorado State University


Department of Biology
Colorado State University
1878 Campus Delivery
Fort Collins, CO 80523


  • Professor, Department of Biology, Colorado State University


  • MS, Botany, University of Wyoming-Laramie, 1981
  • BS, Biology (high honors), Idaho State University, 1978
  • PhD, Botany, University of Wyoming-Laramie

External Links

Journal Articles


Knapp, A. K., M. L. Avolio, C. Beier, C. J. Carroll, S. L. Collins, J. S. Dukes, L. H. Fraser, R. J. Griffin-Nolan, D. L. Hoover, A. Jentsch, M. E. Loik, R. P. Phillips, A. K. Post, O. E. Sala, I. J. Slette, L. Yahdjian and M. D. Smith. 2017. Pushing precipitation to the extremes in distributed experiments: Recommendations for simulating wet and dry years. Global Change Biology 23(51774-1782):. DOI: 10.1111/gcb.13504. (link )

Wilcox, K. R., Z. Shi, L. Gherardi Arbizu, N. P. Lemoine, S. E. Koerner, D. L. Hoover, E. W. Bork, K. M. Byrne, J. F. Cahill Jr, S. L. Collins, S. Evans, A. K. Gilgen, P. Holub, L. Jiang, A. K. Knapp, D. LeCain, J. Liang, P. Garcia-Palacios, J. Penuelas, W. T. Pockman, M. D. Smith, S. Sun, S. R. White, L. Yahdjian, K. Zhu and Y. Luo. 2017. Asymmetric responses of primary productivity to precipitation extremes: A synthesis of grassland precipitation manipulation experiments. Global Change Biology 23(10):4376-4385. DOI: 10.1111/gcb.13706. (link )

Wilcox, K. R., A. T. Tredennick, S. E. Koerner, E. Grman, L. M. Hallett, M. L. Avolio, K. J. La Pierre, G. R. Houseman, F. isbell, D. S. Johnson, J. M. Alatalo, A. H. Baldwin, E. W. Bork, E. H. Boughton, W. D. Bowman, A. J. Britton, J. F. Cahill Jr, S. L. Collins, G. Du, A. Eskelinen, L. Gough, A. Jentsch, C. Kern, K. Klanderud, A. K. Knapp, J. Kreyling, Y. Luo, J. R. McLaren, P. Megonigal, V. G. Onipchenko, J. Prevey, J. N. Price, C. H. Robinson, O. E. Sala, M. D. Smith, N. A. Soudzilovskaia, L. Souza, D. Tilman, S. R. White, Z. Xu, L. Yahdjian, Q. Yu, P. Zhang and Y. Zhang. 2017. Asynchrony among local communities stabilizes ecosystem function of meta-communities. Ecology Letters 20(12):1534-1545. DOI: 10.1111/ele.12861. (link )


Knapp, A. K., D. L. Hoover, K. R. Wilcox, M. L. Avolio, S. E. Koerner, K. J. La Pierre, M. E. Loik, O. E. Sala and M. D. Smith. 2015. Characterizing differences in precipitation regimes of extreme wet and dry years: Implications for climate change experiments. Global Change Biology 21(7):2624-2633. DOI: 10.1111/gcb.12888. (link )


Levy, O., B. A. Ball, B. Bond-Lamberty, K. S. Cheruvelil, A. O. Finley, N. R. Lottig, S. W. Punyasena, J. Xiao, J. Zhou, L. B. Buckley, C. T. Filstrup, T. H. Keitt, J. R. Kellner, A. K. Knapp, A. D. Richardson, D. Tcheng, M. Toomey, R. Vargas, J. W. Voordeckers, T. Wagner and J. W. Williams. 2014. Approaches to advance scientific understanding of macrosystems ecology. Frontiers in Ecology and the Environment 12(1):15-23. DOI: 10.1890/130019. (link )


Collins, S. L., S. R. Carpenter, S. M. Swinton, D. E. Orenstein, D. L. Childers, T. L. Gragson, N. B. Grimm, J. M. Grove, S. L. Harlan, J. P. Kaye, A. K. Knapp, G. P. Kofinas, J. J. Magnuson, W. H. McDowell, J. M. Melack, L. A. Ogden, G. P. Robertson, M. D. Smith and A. C. Whitmer. 2011. An integrated conceptual framework for social-ecological research. Frontiers in Ecology and the Environment 9(6):351-357. (link )

Knapp, A. K., M. Mincarone, H. Harwell, B. A. Polidoro, J. C. Sanciangco and K. E. Carpenter. 2011. Conservation status of the world's hagfish species and the loss of phylogenetic diversity and ecosystem function. Aquatic Conservation: Marine and Freshewater Ecosystems 21(5):401-411. DOI: 10.1002/aqc.1202. (link )


Foster, D. R., F. Swanson, J. D. Aber, J. Burke, N. Brokaw, D. Tilman and A. K. Knapp. 2003. The importance of land-use legacies to ecology and conservation. BioScience 53(1):77-88.



Knapp, A. K., J. M. Briggs, D. C. Harnett and S. L. Collins. 1998. Grassland dynamics: Long-term ecological research in Tallgrass Prairies. Oxford University Press. Oxford and New York.

Book Chapters


Knapp, A. K., J. M. Briggs, D. L. Childers and O. E. Sala. 2007. Estimating aboveground net primary production in grassland- and herbaceous-dominated ecosystems. Pp. 27-48 In: Fahey, T. J. and A. K. Knapp eds., Principles and Standards for Measuring Primary Production (Long-Term Ecological Research Network). Oxford University Press.



Knapp, A. K., B. A. Polidoro, J. J. Alvarado, M. Benavides, K. E. Carpenter, C. Conand, R. U. Gamboa, J. Hamel, H. Harwell, A. Mercier, E. P. Ortiz, S. Purcell and M. V. Toral-Granda. 2012. Extinction risk of the world's sea cucumber species (order Aspidochirotida). Presentation at the 22-24 March 2012 Benthic Ecology Meeting, Norfolk, VA.

Report Chapters


Yao, J., O. E. Sala and D. P. Peters. 2013. Cross-site studies "by design": Experiments and observations that provide new insights. Pp. 72-80 In: Peters, D. P., C. M. Laney, A. E. Lugo, S. L. Collins, C. T. Driscoll, P. M. Groffman, J. M. Grove, A. K. Knapp, T. K. Kratz, M. D. Ohman, R. B. Waide and J. Yao eds., Long-Term Trends in Ecological Systems: A Basis for Understanding Responses to Global Change. U.S. Department of Agriculture. Technical Bulletin 1931. Washington, D.C. (link )